According to your config, UDP/7778 would be required for chat channel/in-game mail, but that would not stop players zoning-in and playing, just stop chat-channel/in-game mail from working.
UDP/9000 (world.exe) and UDP/7000-7100 (zones) should be all that you need for getting into a zone. (actually UDP/7000 to 7004 if you are only booting 5 zone servers).
Something is stopping UDP/700x, but since you say your firewall is off and ports are forwarded, I don't know what it could be.
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